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Steam Deal: Save 90 % on METRO 2033 Redux (1,99€ | Next Gen Update now available)
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Mostly the atmosphere, the environment in the original is a lot more otherworldly like S.T.A.L.K.E.R. and they removed pretty much all of that. They replaced moody art direction with more "photorealistic" graphics (at the time) and I don't think they hold up as well - this video shows it off some. The mutants are also scarier looking and more varied and it had much more of a horror vibe that was made more into action-adventure, and that applies to the combat too.
I did just find that it is actually available for sale again on gog and for only 1 USD more than this sale, so that's good to see. I still own the original release on steam.